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Before Thunderbird can display the message body, it must download it from the email server.The only difference is that rather than downloading the entire message, only the message "headers" are downloaded.(Message headers contain information like the sender, recipient(s), subject, etc - everything except the body of the message.) The message body is not downloaded until you click on the message in the message list (which means that if you are not online, you can't read the message).You can set general synchronization preferences (such as synchronize recent messages or messages under a certain size).
This part of the page also has options for batch-deleting messages (which should, of course, be used with caution).
Synchronization impacts performance and disk space usage.